Research support was provided through the projects Forest Health Assessment of Cooperative Agreement SRS 05-CA-11330146-180 and Forest Health Monitoring, Analysis and Assessment of Research Joint Venture Agreement 06-JV-11330146-124 between the U. S. Forest Service and North Carolina State University (in accordance with U.S. Department of Agriculture policy, North Carolina State University is prohibited from discriminating on the basis of race, color, national origin, sex, age, or disability. [Not all prohibited bases apply to all programs.]) This research was supported by funds provided by the U.S. Forest Service, Southern Research Station, Asheville, NC and the U.S. Forest Service National Forest Health Monitoring Program. The authors thank the following reviewers for their useful comments: William Bechtold (Southern Research Station), Borys Tkacz (Forest Health Protection), Robert Mangold (Forest Health Protection) and James Steinman (Forest Health Protection).

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